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Did you know paper has a grain? The best analogy I've heard to date is, imagine logs floating down a river. The logs will point with the river's flow. This is exactly what happens with paper when it's made. While there will always been crossing fibers, the bulk of the fiber lays itself pointing in one direction; this is the grain direction.
Sheets are considered to be either long grain or short grain. This refers to the edge that the grain runs parallel with. A 12x18 sheet would be long grain; the grain direction runs parallel to the 18" (long) side of the sheet. An 18x12 sheet would be short grain; where the grain direction runs parallel to the 12" (short) side. Grain direction is often indicated by the second number of the sheet size.
Why is this important? Scoring, folding and binding quality is highly dependent on the grain direction. For the most part, this isn't something you--the designer--has to worry about when ordering things such as booklets or greeting cards with Fresh Impression. This is something taken into account for you behind the scenes, but if you take advantage of ordering full press sheets, it's important to keep grain direction in mind. Be sure to layout your major scores parallel to the grain direction, and indicate the short or long grain direction for the sheet. . .